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Zanthoxylum dipetalum var. dipetalum

This name is accepted

Kingdom: Viridiplantae Phylum: Magnoliophyta Class/Clade: Eudicot-Rosids Order: Sapindales Family: Rutaceae Genus: Zanthoxylum

kāwa‘u, kāwa‘u kua kuku kapa

Description

Key Characters:

Growth Form: Trees 4–15 m tall.

Stems: Trunk up to 30 cm in diameter, bark finely fissured–reticulate, new growth glabrous or finely puberulent to tomentose, young branches subtended by a pair of bud scales.

Roots:

Leaves: Leaves compound. Alternate. Leaflets 3–7, thick, coriaceous, lowest pair strongly reduced, (0.2–)1.5–3(–4) cm long, rarely 1 or both reduced leaflets absent, normal leaflets 6–15(–36) cm long, (2.5–)3.5–9(–13.5) cm wide, lateral petiolules articulated only at base, 3–9 mm long, terminal petiolule twice articulated, at base and near leaflet, 8–30 mm long; lowest pair of leaflets stipelliform or, if expanded, then usually folded or strongly revolute, arising immediately below lowest pair of normal leaflets, normal leaflets oblong–elliptic, elliptic to broadly elliptic or sometimes broadly obovate. Leaflet apices obtuse to rounded. Base cuneate to rounded. Surfaces glabrous; conspicuously glandular punctate. Margins entire rarely weakly emarginate. Normal leaflets lateral veins usually 6–8 pairs, prominent, higher order venation forming a reticulate pattern. Rachis terete (cylindrical) or winged. Stipules absent.

Flowers: Flowers usually 5–15 in terminal or leaf–opposed, open, cymose inflorescences usually 5–10 cm long, peduncles 10–40 mm long, pedicels 3–6(–8) mm long, subtended by a pair of bud scales; bracteoles 1 mm or less long. Flowers unisexual, actinomorphic. Calyx of 4 sepals, broadly ovate–deltate, 1–1.5 mm long, ciliolate. Corolla of 2 or sometimes 4 petals; petals white or yellowish white, sometimes tinged reddish, lanceolate to ovate, 6–10 mm long. Stamens 4, alternate with the petals, glabrous or pubescent, absent in pistillate flowers; anthers medifixed, dithecal. Ovary superior, 2-carpellate, but usually only 1 carpel developing, placentation axile, rudimentary in staminate flowers; ovules 2 per carpel, pendulous; style short, ± coherent; stigmas obliquely orbicular, capitate.

Fruit: Follicles 1–2; ovoid; 15–25(–33) mm long; glandular–pitted; apex conspicuously beaked; the beak 3–8 mm long; exocarp rugose; endocarp coriaceous. Seeds usually 1 per follicle; on a slender funicle; black; ovoid; 10–26 mm long.

Ploidy:

Habitat: Dry to mesic forest; sometimes along margins of wet forest.

Elevation Range: 260–1,280 m.

Historical Distribution

Uses and Culture

USES

Natural History

Statewide Status

Endemic

Island Status

Kaua'i Endemic
O'ahu Endemic
Molokai Endemic
Maui Endemic
Hawai'i Endemic

Dispersal Agents


Pollinators

Bibliography

Other References

Wagner et al. 1990:1214 (K, O, Mo, H); Faccenda, Arthur, et al. 2025:210 (WM); Hank Oppenheimer pers. obs. (Mo†, 1985)

Occurrences

SNo. Scientific Name Locality Habitat Basis of Record Recorded By Record Number Island Source Date
1 Zanthoxylum dipetalum var. dipetalum Honopu Trail Trail-side Preserved_Specimen Marr, K.L. 40 Kauai BISH 1988-06-11
2 Zanthoxylum dipetalum var. dipetalum Kawaikoi Stream, by Sugi grove Along bank of stream. Preserved_Specimen Corn, C. Kauai BISH 1982-08-29
3 Zanthoxylum dipetalum var. dipetalum Makaha Rd, 1.5 m down valley, in sm valley N of Kakio Keakeo Picnic area N facing slope Preserved_Specimen Marr, K.L. 12 Kauai BISH 1988-03-26
4 Zanthoxylum dipetalum var. dipetalum Lihue-Koloa FR, ridge ESE of Puu Kolo Metrosideros, Acacia, Canthium, Cyanea, Bidens, Myrsine, Dicranopteris Preserved_Specimen Flynn, T.W. 3149 Kauai BISH 1988-09-21
5 Zanthoxylum dipetalum var. dipetalum 2 ridges S. of Pohakea Pass forest Preserved_Specimen Christophersen, E. 1658 Oahu BISH 1931-03-19
6 Zanthoxylum dipetalum var. dipetalum HVNP, Kipuka Puaulu. Next to NPS transect One other large individual and at least 5 other saplings in Kipuka Puaulu. Three saplings and one seedling beneath this tree. Preserved_Specimen Drake, D. 0.19 Hawaii BISH 1988-03-25
7 Zanthoxylum dipetalum var. dipetalum Kumuwela Ridge, E of Waineke Swamp Forest Preserved_Specimen Degener, O. 21551 Kauai BISH 1952-01-08
8 Zanthoxylum dipetalum var. dipetalum Waimea Canyon, mile 11.3, Hwy. 550 Zanthoxylum kauaense, Pelea anisata, Myrsine, Acacia; dry Preserved_Specimen Flynn, T.W. 2212 Kauai BISH 1987-05-21
9 Zanthoxylum dipetalum var. dipetalum Halemanu Diverse native forest, canopy emergent to 7+ m Preserved_Specimen Takeuchi, W.N. 3678 Kauai BISH 1987-09-26
10 Zanthoxylum dipetalum var. dipetalum Bird Park (Kipuka Puaulu) moderate forest Preserved_Specimen Morley, H. 104 Hawaii BISH 1934-08-11
11 Zanthoxylum dipetalum var. dipetalum Kumuwela Ridge, Waimea Dry open woods Preserved_Specimen St.John, H. 13822 Kauai BISH 1933-12-28
12 Zanthoxylum dipetalum var. dipetalum Puu Kaua, Waianae Mts. Honouliuli wooded steep east ridge Preserved_Specimen St.John, H. 12226 Oahu BISH 1932-11-06
13 Zanthoxylum dipetalum var. dipetalum HVNP, Kipuka Puaulu. Next to NPS transect One other large individual and at least 5 other saplings in Kipuka Puaulu. Three saplings and one seedling beneath this tree. Preserved_Specimen Drake, D. 0.19 Hawaii BISH 1988-03-25
14 Zanthoxylum dipetalum var. dipetalum Waianae Mts., Ekahnanui Gulch, ridge, E Puu Kaua Myrsine emarginata, Myrsine sandwicensis, Exocarpos gaudichaudii, Styphelia, Dubautia, Plantaginea, Pittosporum flocculosum, Scaevola gaudichaudiana Preserved_Specimen Perlman, S. 5100 Oahu BISH 1987-01-15
15 Zanthoxylum dipetalum var. dipetalum ridge above Ekahanui gulch Schinus terebinthifolius Preserved_Specimen Takeuchi, W.N. 2063 Oahu BISH 1985-03-10
16 Zanthoxylum dipetalum var. dipetalum Upper Limahuli Valley down stream from base camp up drainage on W side. Metrosideros-Dicranopteris wet forest. W/ Diospyros sandwicensis, Diplopterygium pinnatum, Bobea elatior, Ilex, Heyotis terminalis, Tetraplasandra, Pittosporum kauaiense, Pouteria, Coprosma, Boehmeria, Antidesma, Broussaisia, Charpentiera Preserved_Specimen Tangalin, N. 898 Kauai BISH 2006-08-08
17 Zanthoxylum dipetalum var. dipetalum Oio, Paumalu Trail forest Preserved_Specimen Degener, O. 17724 Oahu BISH 1940-06-16
18 Zanthoxylum dipetalum var. dipetalum Kokee. App. 30m past Kokee State Park sign on right side of main road. Kokee State Park Preserved_Specimen Marr, K.L. 0.15 Kauai BISH 1988-03-26
19 Zanthoxylum dipetalum var. dipetalum Puu Kaua, Waianae Mts., Honouliuli wooded steep e. ridge Preserved_Specimen St.John, H. 12226 Oahu BISH 1932-11-06
20 Zanthoxylum dipetalum var. dipetalum HVNP, Kipuka Puaulu. Next to NPS transect One other large individual and at least 5 other saplings in Kipuka Puaulu. Three saplings and one seedling beneath this tree. Preserved_Specimen Drake, D. 0.19 Hawaii BISH 1988-03-25
21 Zanthoxylum dipetalum var. dipetalum Waioli-Waipa ridge, 10 ft. e. of crest bse of large boulder Preserved_Specimen Christensen, C. 195 Kauai BISH 1977-04-27
22 Zanthoxylum dipetalum var. dipetalum Kokee, Awaawapuhi trail Forest Preserved_Specimen Stone, B.C. 3338 Kauai BISH 1960-04-12
23 Zanthoxylum dipetalum var. dipetalum Waimea District; Kawaiula Valley, Kuia Natural Area Reserve Metrosideros-Acacia mesic forest with Alphitonia ponderosa, Diospyros sandwicensis, dracaena aurea, Psychotria mariniana, Dodonaea viscosa, Wikstroemia furcata, Planchnella sandwicensis, dryopteris glabra, Doodia kunthiana, Panicum Preserved_Specimen Wood, K.R. 14967 Kauai BISH 2012-05-09
24 Zanthoxylum dipetalum var. dipetalum Hanalei District, Kalalau, valley slopes below Puu o Kila. Metrosideros mixed mesic forest asjacent to mesic cliffs w/ Planchonella sandwicensis, Dodonaea viscosa, Psydrac oderata, Diospyros sandwicensis, Chrysodracon aurea, Santalum pyrularium, Wikstroemia oahuensis, Rauvolfia sandwicensis, Syzygium sandiwcensis Preserved_Specimen Wood, K.R. 18298 Kauai BISH 2019-08-28
25 Zanthoxylum dipetalum var. dipetalum Pohakuao, hanging valley between Kalalau and Hanakoa Lowland mesic forest Preserved_Specimen Wood, K.R. 1769 Kauai BISH 1992-04-02
26 Zanthoxylum dipetalum var. dipetalum Makakupaia, S fork, Kaunakakai Gulch Xylosma, Coprosma, Hedychium, Osmanthus, Wikstroemia, Melinis, Microlepia Preserved_Specimen Lau, J. 1465 Molokai BISH 1985-09-25
27 Zanthoxylum dipetalum var. dipetalum Anahola Mountains, near SE end of range. Mesic forest on ridge. W/ Metrosideros, Dicranopteris, Carex, Chrysodracon, Psydrax, Machaerina mariscoides. Preserved_Specimen Edmonds, M. 309 Kauai BISH 2015-03-22
28 Zanthoxylum dipetalum var. dipetalum Waimea District; Kawaiiki Zone, Below Kalukaula Ridge, on ridge looking down into drainage of Koaie Canyon and north of Waialae. Degraded Metrosideros - Acacia koa Diverse Mesic Forest Myoporum, Cryptocarya, Kadua affinis, Alyxia, Claoxylon sandwicensis, Dodonea, Leptocaphylla, Dryopteris, Zanthoxylum, Dubautia, Psychotria mariniana, Scaevola procera, Melicope Preserved_Specimen Tangalin, N. 2573 Kauai BISH 2011-03-23
29 Zanthoxylum dipetalum var. dipetalum Hind ranch, between Puuwaawaa and Puu Iki Growing in open cattle pasture, with Reynoldsia, Osmanthus, Sophora, Dracaena, Nototrichium, and other dry forest trees. Deep soil phase. Preserved_Specimen Hatheway, W.H. 452 Hawaii BISH 1950-11-20
30 Zanthoxylum dipetalum var. dipetalum Honopu, Awaawapuhi Valley Mesic forest, canopy ht ave 55 ft Preserved_Specimen Stone, B.C. 1592 Kauai BISH 1956-12-29
31 Zanthoxylum dipetalum var. dipetalum Koolau Range, Pupukea-Paumalu Forest Preserve, Pupukea Road moist slope below trail Preserved_Specimen Cowan, R.S. 774 Oahu BISH 1948-01-25
32 Zanthoxylum dipetalum var. dipetalum West Maui, Lahaina District, kauaula Valley, south slope. Vegetation: Pisonia, Myrsine, Diospyros, Metrosideros, Planchonella, Nestegis, Leptecophylla, Hibiscadelphus, Chrysodracon, Dodonaea, Charpentiera, Euphorbia, Myoporum, Santalum, Coprosma, Psychotria, Alyxia, Dryopteris, Ipomoea, Cocculus, Trisetum, Cyrtomium, Microlepia, Carex, Doodia, Bidens, Peperomia, Ageratina, Rubus, Lantana, Oplismenus, Bromus, _rigeron. Preserved_Specimen Oppenheimer, H.L. 41608 Maui BISH 2016-04-13
33 Zanthoxylum dipetalum var. dipetalum Hikimoe, N facing slopes below road Small patches of Diospyros sandwicensis forest with occasional Metrosideros polymorpha, Coprosma waimeae, Psychotria greenwelliae, Zanthoxylum dipetelum, Psydrax odorata, Pouteria sandwicensis, Nestegis sandwicensis, Pleomele aurea, Pisonia sp., Syzygium sandwicensis, gulch is mostly dominated by secondary invasive species. Preserved_Specimen Wood, K.R. 14226 Kauai BISH 2010-06-01
34 Zanthoxylum dipetalum var. dipetalum Kaunala Gulch, Kaunala, Koolau Range, Kahuku Forest Res., lower woods lower woods Preserved_Specimen St.John, H. 20164 Oahu BISH 1941-02-09
35 Zanthoxylum dipetalum var. dipetalum N Kaala, Dupont trail forest Preserved_Specimen Degener, O. 19431 Oahu BISH 1949-02-22
36 Zanthoxylum dipetalum var. dipetalum Oio, Paumalu Trail forest Preserved_Specimen Degener, O. 17590 Oahu BISH 1940-06-16
37 Zanthoxylum dipetalum var. dipetalum Na Pali Survey, Supp. Sta. C., back of valley on E side Diverse mesic forest on steep slope Preserved_Specimen Perlman, S. 10374 Kauai BISH 1989-06-13
38 Zanthoxylum dipetalum var. dipetalum Waianae Mts., S Ekanui Gulch, ridge Metrosideros, Myrsine, Schinus Preserved_Specimen Obata, J.K. 394 Oahu BISH 1978-07-09
39 Zanthoxylum dipetalum var. dipetalum Maui, W Maui, Lahaina District, Kauaula Valley, S slope on steep, rocky slope. Vegetation: Metrosideros, Sophora, Nestegis, Coprosma, Dodonaea, Pipturus, Achyranthes, Diospyros, Myrsine, Melicope, Pleomele, Remya, Dryopteris, Senna, Ipomoea, Neraudia, Lysimachia, Wikstroemia, Osteomeles, Trisetum, Panicum, Microlepia, Lysimachia, Eragrostis, Alyxia, Lipochaeta, Bidens, Peperomia, Ageratina, Lantana, Grevillea, Conyza, Petroselinum. Preserved_Specimen Oppenheimer, H.L. 41338 Maui BISH 2013-04-24
40 Zanthoxylum dipetalum var. dipetalum Awaawapuhi Trail; Honopu Moist Acacia Forest. Preserved_Specimen Lane, I.E. 56625 Kauai BISH 1956-12-29
41 Zanthoxylum dipetalum var. dipetalum Hanalei, Hoolulu Valley Nr ridge crest in mixed mesic forest Preserved_Specimen Lau, J. 2205 Kauai BISH 1986-04-01
42 Zanthoxylum dipetalum var. dipetalum Kamiloloa, ridge above Gulch dry; native trees and shrub; exotic grass and shrubs Preserved_Specimen Stemmermann, R.L. 3919 Molokai BISH 1979-07-15
43 Zanthoxylum dipetalum var. dipetalum Bird Park (Kipuka Puaulu) moderate forest Preserved_Specimen Morley, H. 104 Hawaii BISH 1934-08-11
44 Zanthoxylum dipetalum var. dipetalum Lahaina District. Kauaula valley, South slope, below Helu summit, west Maui Diospyros sandwicensis-Metrosideros mesic forest. With Nestegis sandwicensis, Pisonia, Zanthoxylum hawaiiense, Cyrtandra grayi, Coprosma foliosa, Cheirodendron trigynum, Dodonaea viscosa, Polyscias oahuensis, Myrsine lanaiensis, Sophora chrysophylla Preserved_Specimen Perlman, S. 23854 Maui BISH 2014-02-13
45 Zanthoxylum dipetalum var. dipetalum Kahuku mauka shrubby forest Preserved_Specimen Degener, O. 19544 Oahu BISH 1949-03-20
46 Zanthoxylum dipetalum var. dipetalum Kumuwela Ridge, E of Waineke Swamp Forest Preserved_Specimen Degener, O. 21551 Kauai BISH 1952-01-08
47 Zanthoxylum dipetalum var. dipetalum road to Alakai Swamp Preserved_Specimen MacDaniels, L.H. 764 Kauai BISH 1927-02-17
48 Zanthoxylum dipetalum var. dipetalum Waianae Mts., Honouliuli FR, ridge to Puu Kaua exposed arid cliff Preserved_Specimen Herbst, D.R. 5263 Oahu BISH 1975-03-16
49 Zanthoxylum dipetalum var. dipetalum Honopu Trail Beside trail Preserved_Specimen Marr, K.L. 14 Kauai BISH 1988-03-26
50 Zanthoxylum dipetalum var. dipetalum Kalahu, W of Kalalau Valley Open Dicranopteris forest Preserved_Specimen Degener, O. 20596 Kauai BISH 1950-02-01
51 Zanthoxylum dipetalum var. dipetalum Koloa, Haupu open shrubland, herbland, sparse tree cover; Metrosideros polymorpha, Dicranopteris linearis, Cibotium glaucum, Sadleria pallida, Hedyotis terminalis, Psychotria mariniana, Antidesma platyphyllum, Bobea brevipes, Xylosma hawaiiense, Cheirodendron fauriei, Melicope feddei, Tetraplasandra Preserved_Specimen Wood, K.R. 11973 Kauai BISH 2006-07-14
52 Zanthoxylum dipetalum var. dipetalum Hanalei, Honoonapali NAR, West Hanakoa stream drainage above falls. Metrosideros-Cheirodendron-Dicranopteris wet forest w/ Kadua affinis, Psychotria mariniana, P. mauiensis, Melicope clusiifolia, M. anisata, M. degeneri, Ilex anomala, Syzygium sandwicensis, Elaeocarpus bifidus, Dodonaea viscosa, Santalum involutum, Kad... Preserved_Specimen K.R. Wood 19117 Kauai BISH 2022-07-22
53 Zanthoxylum dipetalum var. dipetalum Halemanu, Kokee Open woods Preserved_Specimen Degener, O. 28099 Kauai BISH 1960-12-02
54 Zanthoxylum dipetalum var. dipetalum Kalalau valley, drainage west of Pohakulele Camp and east of Huna. Diospyros mixed mesic forest occasional in area Preserved_Specimen Wood, K.R. 11655 Kauai BISH 2005-12-15
55 Zanthoxylum dipetalum var. dipetalum Kumuwela Ridge, Waimea Dry open woods Preserved_Specimen St.John, H. 13835 Kauai BISH 1933-12-28
56 Zanthoxylum dipetalum var. dipetalum HVNP, Kipuka Puaulu, next to NPS transect Sapling growing beneath KM 19. Preserved_Specimen Marr, K.L. 91 Hawaii BISH 1988-08-01
57 Zanthoxylum dipetalum var. dipetalum Plateau just S of Awaawapuhi stream head Diverse mesic forest intermixed with open patches of Dicranopteris. With Metrosideros, Ilex, Scaevola sp., Dodonaea, Pouteria, Psychotria greenwelliae, Melicope anisata, Coprosma, Psidium cattleianum, Carex meyenii, Alyxia stellata, and occasional Hedychium. Preserved_Specimen Trauernicht, C. 750 Kauai BISH 2009-02-19
58 Zanthoxylum dipetalum var. dipetalum N Haupu moist Preserved_Specimen MacDaniels, L.H. 737 Kauai BISH 1927-02-16
59 Zanthoxylum dipetalum var. dipetalum Kohua Ridge Forest Preserved_Specimen Degener, O. 23920 Kauai BISH 1955-09-09
60 Zanthoxylum dipetalum var. dipetalum Halemanu Dry-mesic forest. Preserved_Specimen Herbst, D.R. 1030 Kauai BISH 1968-04-07
61 Zanthoxylum dipetalum var. dipetalum Puu Kaua, S Ekahanui, subgulch Alectryon, Elaeocarpus, Pouteria, Antidesma, Pisonia, Charpentiera, Melicope, Freycinetia, Claoxylon, Hibiscus, Pipturus, Passiflora suberosa, Labordia, Platydesma, Athyrium sandwichianum, Neraudia Preserved_Specimen Perlman, S. 6113 Oahu BISH 1987-06-28
62 Zanthoxylum dipetalum var. dipetalum HVNP, Kipuka Puaulu, next to NPS transect Sapling growing beneath KM 19. Preserved_Specimen Marr, K.L. 91 Hawaii BISH 1988-08-01
63 Zanthoxylum dipetalum var. dipetalum Pupukea Paumala FR, Koolau Mts., jeep road ohia, Eugenia Preserved_Specimen Spence, G. 8 Oahu BISH 1971-10-31
64 Zanthoxylum dipetalum var. dipetalum Puu Kaua, E ridge Exocarpos; Myrsine, Dubautia, Pelea, Metrosideros Preserved_Specimen Marr, K.L. 52 Oahu BISH 1988-06-24
65 Zanthoxylum dipetalum var. dipetalum Waimea district Kawaiiki ridge, slope of Koaie, west of twin falls Metrosideros- Acacia montane mixed mesic forest, summer dry Diospyros sandwicensis, Pleomele aurea, Myrsine lanaiensis, Planchonella sandwicensis, Cryptocarya manii, Myoporum sandwicensis, Nestegis sandwicensis, Psydrax odorata, Zanthoxylum dipetalum, Pteraly Preserved_Specimen Wood, K.R. 15418 Kauai BISH 2013-02-17
66 Zanthoxylum dipetalum var. dipetalum ridge, Puu Kaua cliff edge Preserved_Specimen Takeuchi, W.N. 126 Oahu BISH 1984-07-15
67 Zanthoxylum dipetalum var. dipetalum E of Kumuwela Ridge, Kokee Open forest along ditch trail Preserved_Specimen Degener, O. 20598 Kauai BISH 1950-02-07
68 Zanthoxylum dipetalum var. dipetalum Ridge .75 mi N of Laaukahi, Koloa Wooded NW-facing slope Preserved_Specimen St.John, H. 23011 Kauai BISH 1947-12-24
69 Zanthoxylum dipetalum var. dipetalum Waimea, 1.3 mi. down Makaha Ridge road from Hwy. 550, below Kokio kookeo picnic area Acacia koa, Metrosideros, Pelea barbigera, Remya kauaiensis, Alphitonia Preserved_Specimen Flynn, T.W. 2773 Kauai BISH 1988-02-24
70 Zanthoxylum dipetalum var. dipetalum ridge, Puu Kaua, Honouliuli FR Metrosideros, Acacia koa, grass Preserved_Specimen Nagata, K.M. 1156 Oahu BISH 1973-06-11
71 Zanthoxylum dipetalum var. dipetalum Kumuwela Ridge, Waimea Dry open woods Preserved_Specimen St.John, H. 13835 Kauai BISH 1933-12-28
72 Zanthoxylum dipetalum var. dipetalum Honopu Trail Beside trail Preserved_Specimen Marr, K.L. 41 Kauai BISH 1988-06-11
73 Zanthoxylum dipetalum var. dipetalum ridge, Kaua to Kanehoa Psychotria hathewayi Preserved_Specimen Takeuchi, W.N. Oahu BISH 1986-10-01
74 Zanthoxylum dipetalum var. dipetalum Kalalau Valley, S rim and upper reaches, 0.4 miles E of Kalalau Lookout; steep slopes with low mixed mesophytic forest of Metrosideros, Pelea, Xylosma & Pleomele Preserved_Specimen Lorence, D.H. 5121 Kauai BISH 1987-03-20
75 Zanthoxylum dipetalum var. dipetalum Kalalau, very back on E side of Valley, 50 ft up talus slope on E side of N running gulch, gulch just W of the mainstream. Old relic mixed mesic forest with weedy understory. Christella parasitica, Nephrolepis, Bryophyllum pinnatum, Adenophorus hispidulum, Elephantopus mollis, midstory mostly open with Lantana and Psydrax, loose open canopy of Aleurites moluccana, Melia azedarach Preserved_Specimen Tangalin, N. 2385 Kauai BISH 2010-08-22
76 Zanthoxylum dipetalum var. dipetalum Kumuwela Ridge Forest Preserved_Specimen Degener, O. 28094 Kauai BISH 1955-09-14
77 Zanthoxylum dipetalum var. dipetalum Waimea District. Nuololo, north facing slopes above northern branch. Metrosideros-Acacia montane mesic forest with Alphitonia, Pouteria, Xylosma hawaiiensis, Claoxylon, Myrsine lanaiensis, Wikstromia furcta, Dodonea, Kadua affins, Melicope avata, M. anisate, Dracaena aurea, Polyscias kavaiensis, Psychotri Preserved_Specimen Wood, K.R. 15098 Kauai BISH 2012-07-18
78 Zanthoxylum dipetalum var. dipetalum Kilauea, Kipuka Puaulu Preserved_Specimen Rock, J.F.C. 5916 Hawaii BISH 1914-12-01
79 Zanthoxylum dipetalum var. dipetalum W Waimea drainage basin Preserved_Specimen Forbes, C.N. 1098 Kauai BISH 1917-07-03
80 Zanthoxylum dipetalum var. dipetalum Pupukea Military Trail Preserved_Specimen MacDaniels, L.H. 530 Oahu BISH 1927-01-29
81 Zanthoxylum dipetalum var. dipetalum Awaawapuhi Trail Preserved_Specimen Corn, C. 274 Kauai BISH 1987-06-16
82 Zanthoxylum dipetalum var. dipetalum 2 mi E of Kokee St[ream]. Preserved_Specimen Henrickson, J.S. 4056 Kauai BISH 1969-07-01
83 Zanthoxylum dipetalum var. dipetalum Kawaikoa Stream trail, above trail on S side of stream 70 m before .5 mi marker Preserved_Specimen Marr, K.L. 46 Kauai BISH 1988-06-15
84 Zanthoxylum dipetalum var. dipetalum Kipuka Puaulu Preserved_Specimen Rock, J.F.C. 17152 Hawaii BISH 1918-07-01
85 Zanthoxylum dipetalum var. dipetalum Kaholuamano, near mountain house Preserved_Specimen Rock, J.F.C. 5540 Kauai BISH 1909-09-28
86 Zanthoxylum dipetalum var. dipetalum Mohihi Rd. Kauaikinana Stream, ~20 yds. below dam Preserved_Specimen Flynn, T.W. 438 Kauai BISH 1983-06-02
87 Zanthoxylum dipetalum var. dipetalum Puaulu Preserved_Specimen Rock, J.F.C. 10208 Hawaii BISH 1911-07-01
88 Zanthoxylum dipetalum var. dipetalum Honopu Trail Preserved_Specimen Marr, K.L. 39 Kauai BISH 1988-06-11
89 Zanthoxylum dipetalum var. dipetalum Pupukea-Kahuku Road, Kahuku Preserved_Specimen St.John, H. 25972 Oahu BISH 1956-10-28
90 Zanthoxylum dipetalum var. dipetalum Puu Kaua, Waianae Mts, steep east ridge Preserved_Specimen St.John, H. 20125 Oahu BISH 1940-12-01
91 Zanthoxylum dipetalum var. dipetalum Halemanu Preserved_Specimen Takeuchi, W.N. 3672 Kauai BISH 1987-09-26
92 Zanthoxylum dipetalum var. dipetalum Kauai Preserved_Specimen Knudsen 82 Kauai BISH
93 Zanthoxylum dipetalum var. dipetalum Lava tube region, Kilauea Preserved_Specimen Morley, H. 104 Hawaii BISH 1934-08-06
94 Zanthoxylum dipetalum var. dipetalum Kaholuamano Preserved_Specimen Rock, J.F.C. 10209 Kauai BISH 1911-10-01
95 Zanthoxylum dipetalum var. dipetalum Kaholuamano Preserved_Specimen Rock, J.F.C. 5201 Kauai BISH 1909-09-28
96 Zanthoxylum dipetalum var. dipetalum Puu Kana Preserved_Specimen Topping, D.L. 3286 Oahu BISH 1926-03-21
97 Zanthoxylum dipetalum var. dipetalum Halemanu-Kokee trail, .5 mi fm upper parking lot. Upper end of gully, s sside of trail 20 m fm trail where it levels off Preserved_Specimen Marr, K.L. 43 Kauai BISH 1988-06-13
98 Zanthoxylum dipetalum var. dipetalum Kauaikinana Stream, just below diversion dam Preserved_Specimen Marr, K.L. 13 Kauai BISH 1988-03-26
99 Zanthoxylum dipetalum var. dipetalum Kokee, Milolii Rd, 0.35 mi after jct. with Makaha Rd Preserved_Specimen Marr, K.L. 18 Kauai BISH 1988-03-26
100 Zanthoxylum dipetalum var. dipetalum Kokee, Awaawapuhi trail Preserved_Specimen Stone, B.C. 3337 Kauai BISH 1960-04-12
101 Zanthoxylum dipetalum var. dipetalum Kipuka Puaulu Preserved_Specimen Rock, J.F.C. 13019 Hawaii BISH 1917-08-01
102 Zanthoxylum dipetalum var. dipetalum ~30 m past Kokee sign, main road Preserved_Specimen Marr, K.L. 15 Kauai BISH 1988-03-26
103 Zanthoxylum dipetalum var. dipetalum Kokee, Halemanu Rd, 30 m past turnoff at Kokee State Park sign Preserved_Specimen Marr, K.L. 42 Kauai BISH 1988-06-13
104 Zanthoxylum dipetalum var. dipetalum Waianae Range. South Palawai Gulch Preserved_Specimen Wilbur, R.L. 620 Oahu BISH 1948-03-28
105 Zanthoxylum dipetalum var. dipetalum Kipuka Puaulu Preserved_Specimen Rock, J.F.C. 12572 Hawaii BISH 1915-07-01
106 Zanthoxylum dipetalum var. dipetalum Kilauea Volcano, Kipuka Puaulu Preserved_Specimen Rock, J.F.C. 10208 Hawaii BISH 1911-07-01
107 Zanthoxylum dipetalum var. dipetalum W Waimea drainage basin Preserved_Specimen Forbes, C.N. 970 Kauai BISH 1917-07-03
108 Zanthoxylum dipetalum var. dipetalum Waianae Range, Puu Kaala, above Waialua on the Duponte ridge trail at 2800 ft. alt Preserved_Specimen Stone, B.C. 3185 Oahu BISH 1959-12-20
109 Zanthoxylum dipetalum var. dipetalum Kaholuamano Preserved_Specimen Rock, J.F.C. 2115 Kauai BISH 1909-03-03
110 Zanthoxylum dipetalum var. dipetalum Pupukea - Kahuku Trail, Paunalu - Waimea Ridge. Koolau Range Preserved_Specimen St.John, H. 11585 Oahu BISH 1932-03-06
111 Zanthoxylum dipetalum var. dipetalum Kalalau, on a ridge at the back of the valley Preserved_Specimen Hobdy, R.W. 1815 Kauai BISH 1983-08-20
112 Zanthoxylum dipetalum var. dipetalum Kilauea Volcano, Kipuka Puaulu Preserved_Specimen Rock, J.F.C. 10208 Hawaii BISH 1911-07-01
113 Zanthoxylum dipetalum var. dipetalum Kokee, Awaawapuhi trail, ~40 yds from main trail, at #37 Preserved_Specimen Flynn, T.W. 395 Kauai BISH 1983-04-19
114 Zanthoxylum dipetalum var. dipetalum Kaholuamano Preserved_Specimen Rock, J.F.C. 5201 Kauai BISH 1909-09-28
115 Zanthoxylum dipetalum var. dipetalum Koolau Mts., Pupukea-Kahuku trail Preserved_Specimen Selling, O. 3359 Oahu BISH 1938-09-02
116 Zanthoxylum dipetalum var. dipetalum Waimea Canyon rim, 11.8 mi fm Waimea, 50 m below road Preserved_Specimen Marr, K.L. 44 Kauai BISH 1988-06-13
117 Zanthoxylum dipetalum var. dipetalum Kipuka Puaulu Preserved_Specimen Rock, J.F.C. 12572 Hawaii BISH 1915-07-01
118 Zanthoxylum dipetalum var. dipetalum Kaholuamanu, mountains Preserved_Specimen Forbes, C.N. 347 Kauai BISH 1909-09-01
119 Zanthoxylum dipetalum var. dipetalum Waianae Mts., Puu Kaua to Pohakea Pass, 3rd ridge from summit, NE facing Preserved_Specimen Marr, K.L. 54 Oahu BISH 1988-06-24
120 Zanthoxylum dipetalum var. dipetalum Kaholuamano Preserved_Specimen Rock, J.F.C. 2106 Kauai BISH 1909-03-03
121 Zanthoxylum dipetalum var. dipetalum Waimea, Kauai Preserved_Specimen Mann, H. 581 Kauai BISH 1864-01-01
122 Zanthoxylum dipetalum var. dipetalum Kawaikoa Stream trail, above trail on S side of stream 70 m before .5 mi marker Preserved_Specimen Marr, K.L. 45 Kauai BISH 1988-06-15
123 Zanthoxylum dipetalum var. dipetalum Waimea Canyon rim, below road, 11.8 mi fm Waimea Preserved_Specimen Marr, K.L. 9 Kauai BISH 1988-03-26
124 Zanthoxylum dipetalum var. dipetalum Waimea Canyon, 2 mi. E of Kokee ranger station, road to Molilii [possibly Mohihi Stream], 1st stream crossing where flume begins Preserved_Specimen Henrickson, J.S. 4056 Kauai BISH 1969-08-01
125 Zanthoxylum dipetalum var. dipetalum 3 mi down Makaha ridge road, N facing slope Preserved_Specimen Flynn, T.W. 1810 Kauai BISH 1986-07-15
126 Zanthoxylum dipetalum var. dipetalum Waimea Canyon rim, below road, 11.8 mi fm Waimea Preserved_Specimen Marr, K.L. 10 Kauai BISH 1988-03-26
127 Zanthoxylum dipetalum var. dipetalum Pupukea Trail Preserved_Specimen Swezey, O.H. Oahu BISH 1935-06-16
128 Zanthoxylum dipetalum var. dipetalum Kawaihae uka Preserved_Specimen Hillebrand, W.B. Hawaii BISH
129 Zanthoxylum dipetalum var. dipetalum Puu Kaua, ridge, about 2200 ft. Preserved_Specimen Suehiro, A. Oahu BISH 1932-11-06
130 Zanthoxylum dipetalum var. dipetalum Nuuanu Preserved_Specimen Hillebrand, W.B. Oahu BISH
131 Zanthoxylum dipetalum var. dipetalum Hii Mts Preserved_Specimen Lydgate, J.M. Kauai BISH
132 Zanthoxylum dipetalum var. dipetalum Kipuka Puaulu Preserved_Specimen Rock, J.F.C. Hawaii BISH 1917-08-01
133 Zanthoxylum dipetalum var. dipetalum Kipuka Puaulu Preserved_Specimen Russ, G.W. Hawaii BISH 1930-06-01
134 Zanthoxylum dipetalum var. dipetalum Kawaihae uka Preserved_Specimen Hillebrand, W.B. Hawaii BISH
135 Zanthoxylum dipetalum var. dipetalum Olokele (?) Valley Preserved_Specimen Lydgate, J.M. Kauai BISH 1911-11-01
136 Zanthoxylum dipetalum var. dipetalum Pupukea Paumalu FR, off jeep road Preserved_Specimen Landgraf, M.F. Oahu BISH 1967-05-18
137 Zanthoxylum dipetalum var. dipetalum W Kaunala Preserved_Specimen Russ, G.W. Oahu BISH 1933-05-24
138 Zanthoxylum dipetalum var. dipetalum Pupukea Paumalu FR, off jeep road Preserved_Specimen Landgraf, M.F. Oahu BISH 1967-05-18
139 Zanthoxylum dipetalum var. dipetalum W Kaunala Preserved_Specimen Russ, G.W. Oahu BISH 1933-05-24
140 Zanthoxylum dipetalum var. dipetalum Nuuanu Valley Preserved_Specimen Hillebrand, W.B. Oahu BISH
141 Zanthoxylum dipetalum var. dipetalum Kawaihae uka Preserved_Specimen Hillebrand, W.B. Hawaii BISH
142 Zanthoxylum dipetalum var. dipetalum Kawaihae uka Preserved_Specimen Hillebrand, W.B. Hawaii BISH
143 Zanthoxylum dipetalum var. dipetalum Kilauea Volcano, Kipuka Puaulu Preserved_Specimen Rock, J.F.C. 10208 Hawaii BISH 1911-07-01
144 Zanthoxylum dipetalum var. dipetalum Koolau Range, Pupukea-Kahuku trail, in small ravine near road Preserved_Specimen Pearsall, G.A. 522 Oahu BISH 1960-05-18
145 Zanthoxylum dipetalum var. dipetalum W Kaunala Preserved_Specimen Russ, G.W. 706657 Oahu BISH 1933-05-24
146 Zanthoxylum dipetalum var. dipetalum Oahu Preserved_Specimen St.John, H. 11585 Oahu BISH 1932-03-06
147 Zanthoxylum dipetalum var. dipetalum Haw. Natl. Park Preserved_Specimen Fagerlund, G.O. 827 Hawaii BISH 1943-08-01
148 Zanthoxylum dipetalum var. dipetalum Kilauea, Kipuka Puaulu Preserved_Specimen Rock, J.F.C. 5916 Hawaii BISH 1914-12-01
149 Zanthoxylum dipetalum var. dipetalum Kaholuamano Preserved_Specimen Rock, J.F.C. 10209 Kauai BISH 1911-10-01
150 Zanthoxylum dipetalum var. dipetalum Kilauea Volcano, Kipuka Puaulu Preserved_Specimen Rock, J.F.C. 10208 Hawaii BISH 1911-07-01
151 Zanthoxylum dipetalum var. dipetalum Puu Kaua, Waianae Mts., steep e. ridge Preserved_Specimen St.John, H. 20125 Oahu BISH 1940-12-01
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